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All of the flowers now raised here have developed from those _____ in the forest.    

  A. once they grew             B. they grew once 

  C. that once grew              D. once grew

C


解析:

考查定语从句的关系词的用法。分析句意和句子结构可知those之后是个定语从句,those相当于those flowers,是先行词,因为是生长在森林里的(野)花,grew和关系代词之间是主谓关系,因此that不能省。once在此表示“过去曾”,常与过去时连用,还可表示“每次”,引导从句时表示条件,翻译成“一旦”。

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